Raos Vassilis
March 21, 2025 2025-03-21 16:01Raos Vassilis

Raos Vassilis
Professor of Physiology
Medical School, University of Crete, P.C. 71003 Heraklion, Crete
Professor of Physiology, School of Medicine, University of Crete
● Director, Lab of Movement Physiology
● Director, Interdisciplinary Graduate Programme in the “BRAIN and MIND” Sciences
● Collaborating Researcher, Group of Computational Neuroscience, IACM, FORTH.
Present Academic Address:
Lab of Movement Physiology
Dept of Basic Medical Sciences, School of Medicine, University of Crete
University Campus Voutes, 700 13 Heraklion, Crete, Greece
tel: +30 2810 394512, fax: +30 2810 542115
PROFESSIONAL HISTORY
• Professor of Physiology, University of Crete, (Φ.Ε.Κ. 2338/01-10-2021 τ.Γ).
• Associate Professor of Physiology, University of Crete, (Φ.Ε.Κ. 519/08-06-2017 τ. Γ).
• Assistant Professor of Physiology (tenured), University of Crete, (Φ.Ε.Κ. 998/14-12-2009 τ. Γ).
• Assistant Professor of Physiology, University of Crete, (Φ.Ε.Κ. 201/17-08-2005 τ. ΝΠΔΔ).
• Full Faculty Member, Interdisciplinary Graduate Programme in the “BRAIN and MIND” Sciences, 2003-
• Collaborating Researcher, Institute of Applied and Computational Mathematics, FORTH, 2002-
• Postdoctoral Research Fellow, University of Crete, 1999-2005
• Postdoctoral Research Fellow, University of Parma, 1996-1999
• Research Fellow, University of Verona, 1991
• 6/1989-12/1994: Graduate student, Department of Basic Medical Sciences, School of Medicine, University of Crete, (12/1994 Doctorate).
• 9/1984-5/1989: Undergraduate student, Faculty of Biology, University of Patras (5/1989 Bachelor’s in biology).
• 14/10-20/11/2020: EUPRIM-Net & German Primate Center, Laboratory Animal Science Course on Non-Human Primates (FELASA accredited Course 057/17_48_2020)
• Το disentangle the role of the dorsal and ventral premotor cortex in the control of hand-grasping actions. In particular, to assess the intra- and inter-areal dynamics and examine the causality in the premotor grasping network.
• To reveal how the brain processes observed actions and the mechanisms whereby we assign meaning to the actions of other subjects, by investigating the underlying neural dynamics at the micro, meso, and macroscopic scales.
• To develop decoders for the real-time control of a dexterous hand able to manipulate objects versatilely using neural signals recorded from the forelimb representations of the dorsal and ventral premotor cortical areas.
Peer reviewed publications
Original articles
A1. Raos V, Savaki HE. 2021 Functional Imaging of the Cerebellum during Action Execution and Observation. Cereb Cortex Commun. tgab041, DOI: 10.1093/texcom/tgab041
A2. Papadourakis V, Raos V. 2019. Neurons in the macaque dorsal premotor cortex respond to execution and observation of actions. Cereb Cortex 29(10):4223-4237, DOI: 10.1093/cercor/bhy304.
A3. Papadourakis V, Raos V. 2017. Evidence for the representation of movement kinematics in the discharge of F5 mirror neurons during the observation of transitive and intransitive actions. J Neurophysiol 118(6):3215-3229, DOI: 10.1152/jn.00816.2016.
A4. Raos V, Savaki HE. 2017. The role of the prefrontal cortex in action perception. Cereb Cortex 27(10):4677-4690, DOI: 10.1093/cercor/bhw261.
A5. Raos V, Savaki HE. 2016. Perception of actions performed by external agents presupposes knowledge about the relationship between action and effect. Neuroimage. 132:261-273, DOI: 10.1016/j.neuroimage.2016.02.023.
A6. Kilintari M, Raos V, Savaki HE. 2014. Involvement of the superior temporal cortex in action execution and action observation. J Neurosci. 34(27):8999-9011, DOI: 10.1523/JNEUROSCI.0736-14.2014.
A7. Raos V, Kilintari M, Savaki HE. 2014. Viewing a forelimb induces widespread cortical activations. Neuroimage. 89:122-142, DOI: 10.1016/j.neuroimage.2013.12.010.
A8. Carpaneto J, Raos V, Umiltà MA, Fogassi L, Murata A, Gallese V, Micera S. 2012. Continuous decoding of grasping tasks for a prospective implantable cortical neuroprosthesis. J Neuroeng Rehabil. 9:84, DOI: 10.1186/1743-0003-9-84
A9. Fattori P, Breveglieri R, Raos V, Bosco A, Galletti C. 2012. Vision for action in the macaque medial posterior parietal cortex. J Neurosci. 32(9):3221–3234, DOI: 10.1523/JNEUROSCI.5358-11.2012.
A10. Carpaneto J, Umiltà MA, Fogassi L, Murata A, Gallese V, Micera S, Raos V. 2011. Decoding the activity of grasping neurons recorded from the ventral premotor area F5 of the macaque monkey. Neuroscience. 188:80-94, DOI: 10.1016/j.neuroscience.2011.04.062.
A11. Kilintari M, Raos V, Savaki HE. 2011. Grasping in the Dark Activates Early Visual Cortices. Cereb Cortex. 21(4):949-63, DOI: 10.1093/cercor/bhq175.
A12. Stamos AV, Savaki HE, Raos V. 2010. The spinal substrate of the suppression of action during action observation. J Neurosci. 30:11605-11, DOI: 10.1523/JNEUROSCI.2067-10.2010.
A13. Fattori P, Raos V, Breveglieri R, Marzocchi N, Bosco A, Galletti C. 2010. The dorsomedial pathway is not just for reaching: Grasping neurons in the medial parieto-occipital cortex of the macaque. J. Neurosci. 30:342-9, DOI: 10.1523/JNEUROSCI.3800-09.2010.
A14. Savaki HE, Gregoriou GG, Bakola S, Raos V, Moschovakis AK. 2010. The place code of saccade metrics in the lateral bank of the intraparietal sulcus. J. Neurosci. 30:1118-27, DOI: 10.1523/JNEUROSCI.2268-09.2010.
A15. Evangeliou MN, Raos V, Galletti C, Savaki HE. 2009. Functional Imaging of the Parietal Cortex during Action Execution and Observation. Cereb. Cortex. 19(3):624-39, DOI: 10.1093/cercor/bhn116.
A16. Kattoulas E, Smyrnis N, Mantas A, Evdokimidis I, Raos V, Moschovakis AK. 2008. Arm movement metrics influence saccade metrics when looking and pointing towards a memorized target location. Exp. Brain Res. 189:323-338, DOI: 10.1007/s00221-008-1427-4.
A17. Raos V, Evangeliou MN, Savaki HE. 2007. Mental simulation of action in the service of action perception. J. Neurosci. 27:12675-12683, DOI: 10.1523/JNEUROSCI.2988-07.2007.
A18. Bakola S, Gregoriou GG, Moschovakis AK, Raos V, Savaki HE. 2007. Saccade-related information in the superior temporal motion complex: quantitative functional mapping in the monkey. J. Neurosci. 27:2224-2229, DOI: 10.1523/JNEUROSCI.4224-06.2007.
A19. Raos V, Umilta MA, Murata A, Fogassi L, Gallese V. 2006. Functional properties of grasping-related neurons in the ventral premotor area F5 of the macaque monkey. J. Neurophysiol. 95:709-729, DOI: 10.1152/jn.00463.2005.
A20. Raos V, Umilta MA, Gallese V, Fogassi L. 2004. Functional properties of grasping-related neurons in the dorsal premotor area F2 of the macaque monkey. J. Neurophysiol. 92:1990-2002, DOI: 10.1152/jn.00154.2004.
A21. Raos V, Evangeliou MN, Savaki HE. 2004. Observation of action: grasping with the mind’s hand. Neuroimage. 23:193-201, DOI: 10.1016/j.neuroimage.2004.04.024.
A22. Raos V, Franchi G, Gallese V, Fogassi L. 2003. Somatotopic organization of the lateral part of area F2 (dorsal premotor cortex) of the macaque monkey. J. Neurophysiol. 89:1503-1518, DOI: 10.1152/jn.00661.2002.
A23. Fogassi L, Raos V, Franchi G, Gallese V, Luppino G, Matelli M. 1999. Visual responses in the dorsal premotor area F2 of the macaque monkey. Exp. Brain Res. 128:194-199, DOI: 10.1007/s002210050835.
A24. Murata A, Fadiga L, Fogassi L, Gallese V, Raos V, Rizzolatti G. 1997. Object representation in the ventral premotor cortex (area F5) of the monkey. J. Neurophysiol. 78:2226-2230, DOI: 10.1152/jn.1997.78.4.2226.
A25. Savaki HE, Raos VC, Dalezios Y. 1997. Spatial cortical patterns of metabolic activity in monkeys performing a visually guided reaching task with one forelimb. Neuroscience. 76:1007-1034, DOI: 10.1016/s0306-4522(96)00439-3.
A26. Dalezios Y, Raos VC, Savaki HE. 1996. Metabolic activity pattern in the motor and somatosensory cortex of monkeys performing a visually guided reaching task with one forelimb. Neuroscience. 72:325-333, DOI: 10.1016/0306-4522(95)00516-1.
A27. Raos VC, Dermon CR, Savaki HE. 1995. Functional anatomy of the thalamic centrolateral nucleus as revealed with the [14C]deoxyglucose method following electrical stimulation and electrolytic lesion. Neuroscience. 68:299-313, DOI: 10.1016/0306-4522(95)00114-x.
A28. Raos VC, Savaki HE. 1995. Functional anatomy of the thalamic reticular nucleus as revealed with the [14C]deoxyglucose method following electrical stimulation and electrolytic lesion. Neuroscience. 68:287-297, DOI: 10.1016/0306-4522(95)00113-w.
A29. Raos V, Bentivoglio M. 1993. Crosstalk between the two sides of the thalamus through the reticular nucleus: a retrograde and anterograde tracing study in the rat. J Comp Neurol. 332:145-154, DOI: 10.1002/cne.903320202.
A30. Savaki HE, Raos VC, Dermon CR. 1992. Bilateral cerebral metabolic effects of pharmacological manipulation of the substantia nigra in the rat: unilateral intranigral application of the inhibitory GABAA receptor agonist muscimol. Neuroscience. 50:781-794, DOI: 10.1016/0306-4522(92)90204-f.
A31. Chen S, Raos V, Bentivoglio M. 1992. Connections of the thalamic reticular nucleus with the contralateral thalamus in the rat. Neurosci. Lett. 147:85-88, DOI: 10.1016/0304-3940(92)90780-b.
Review articles
R1. Savaki HE, Raos V. 2019. Action Perception and Motor Imagery: Mental Practice of Action. Prog Neurobiol 175:107-125, DOI: 10.1016/j.pneurobio.2019.01.007.
Conference Proceedings / Book chapters
P1. Ashena N., Papadourakis V., Raos V., Oztop E. 2017. Real-Time Decoding of Arm Kinematics During Grasping Based on F5 Neural Spike Data. In: Cong F., Leung A., Wei Q. (eds) Advances in Neural Networks – ISNN 2017. Lecture Notes in Computer Science, vol 10261, pp 261-268. Springer, Cham. DOI: 10.1007/978-3-319-59072-1_31
• Hellenic Society for Neuroscience (Treasurer, 2019-2021; Secretary, 2022-2023; President, 2024-2025)
• International Brain Research Organisation
• Federation of European Neuroscience Societies
• Society for Neuroscience
• International Neuropsychology Symposium (elected member)